The toning pattern just reflects the surface that the coin was resting on while the toning happened. The raised lines would be die scratches/abrasions. Just happenstance that the toning and die disturbance follow in the same general direction.
Dang! I was hoping someone would have said this was a prototype Die! I can see that it is toning to a degree but I don't think it was from a fingerprint, the lines are too straight and there are raised lines where the toning has taken place. Think I'll soak it and see what's underneath.