Pretty firmly agree with "environmental damage," as in corroded. You are right to question. A fully struck coin and missing both layers of cladding would be quite unusual, I would think, even among missing clad errors.
The cupronickel cladding itself is copper alloyed with 25% nickel. Corrosion reactions might have depleted the nickel at the surface or enriched copper at the surface through replating. It looks like I can see greenish blue colors consistent with corrosion products from copper and nickel. And as others have pointed out, the high spots appear lighter in color, where the discolored surface has worn through into the normal cupronickel cladding.
Not sure if weight is quite the be all end all measurement here, either. The quarter could suffer a great deal of obvious cosmetic damage and still be within tolerance. And while it is most likely that missing clad coins will be light weight due to bonding failure, and maybe have a weaker strike to match, there is a rarer version from a rolling error which doesn't drop the weight according to:
https://www.coinworld.com/news/prec...ly-miss.html