Corrosion (post mint damage). The lineation you see in the metal is probably from the rolling of the nickel plates into strips. Similarly, you see similar stripes from rolling blank strips with woodgrain toning (both sides are always parallel).
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these coins have seen quite a few years in water, with it flowing over them non stop, I'm thinking in a river sitting in a small group of pebble on bedrock. Very similar to what I have detected (nickel coins) in the rivers over the years. The ones at the water line in sand or pebble don't seem to have linear corrosion as do the ones found in the currents. IMO.
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