I think the article is talking about incuse elements on the die, not on the finished coin. It can't be talking about the shield cent because the top paragraph starts in 1950 and the paragraph below the highlights mentions 1970.
The first paragraph in the image talks about Cameo, where a dipping process "produced a light frosting over the entire die. This frosting is also known as a Cameo. The die then was polished with a diamond dust compound. The incused design elements were not affected by this polishing and retained their frosting."
So the frosting was retained on the design elements (lettering, portrait), which is also the "incused design elements" they mention. So the "fine impressions caused by the brush" would then be on the lettering or portrait. That's where you'd have to look.