Apologies for resurecting an old thread, but someone mentioned mint marks. These are also known as initial marks and privy marks. Really the last term is most appropriate, though mint mark is what has stuck*.
Basically they were a form of quality control for hammered coinage. In the early days of English/British coinage the moneyer (person who made the coin) would put their name as part of the legend.
But by Tudor/Stuart times (16th/17th century) most coins were struck in London at the Tower of London. Makes sense really as it's security as a prison and army garrison also made it ideal to hold a mint!
The operation was effectively franchised by the Crown to a Mint Master, who was responsible for ensuring that the quality (weight and purity of metal) of the coins fell within set margins.
The reponsibility for checking this quality was that of the Goldsmiths Company and each year at something called the Trials of the Pyx, a selection of recently made coin was assayed. To allow the Master of the Mint to identify which year's issue was which a special privy (private) mark was incorporated into the design.
Records of the Trials allow us to say when a particular mark was in circulation, so a useful means of identifying a coin when otherwise (apart from in some instances sixpences) they were undated.
This coin for example bears the (R) mark (the mark isn't properly struck up on the reverse) showing it was struck under the authority of Parliament between 15 July 1644 and 12 May 1645 (the King having abandoned London early in the Civil War in 1642, to set up his base in Oxford).

* "Initial mark" isn't strictly correct since the mark generally wasn't an initial, nor did it always appear at the begining of the legend (initially). "Mint mark" is also not quite appropriate because for much of the time there was only one mint (in London), unlike modern 'mint marks' which correctly identify the location of manufacture of a particular coin! Whereas 'privy' marks were only of significance to those who were told about (made privy to) their meaning.